1. Catalan Institute for Water Research (ICRA) Girona 17003 Spain
2. Department of Evolutionary Biology, Ecology and Environmental Sciences Faculty of Biology Universitat de Barcelona (UB) Barcelona 08014 Spain
3. Institute of Aquatic Ecology University of Girona Girona 17003 Spain
4. Department of Environmental Science, Policy, and Management University of California, Berkeley Berkeley California 94720 USA