1. Department of Plant SciencesQuaid‐i‐Azam University Islamabad Pakistan
2. State Key Laboratory of Systematic and Evolutionary Botany, Institute of BotanyChinese Academy of Sciences Beijing China
3. University of Chinese Academy of Sciences Beijing China
4. CAS Key Laboratory of Mountain Ecological Restoration and Bioresource Utilization, Chengdu Institute of BiologyChinese Academy of Sciences Chengdu China
5. State Key Laboratory of Virology, Wuhan Institute of VirologyChinese Academy of Sciences Wuhan China