1. Division of Paleontology; American Museum of Natural History; New York City New York
2. Key Laboratory of Vertebrate Evolution and Human Origin of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences; Beijing China
3. Department of Biology; Indiana University of Pennsylvania; Indiana Pennsylvania
4. Shandong Tianyu Museum of Nature; Pingyi Shandong China
5. Institute of Geology and Paleontology, Linyi University; Linyi Shandong China