1. Laboratoire de Biomathématiques et d’Estimations Forestières Faculté des Sciences Agronomiques Université d’Abomey Calavi Cotonou Benin
2. Plant Ecology and Nature Conservation Group Wageningen University PO Box 47 Wageningen AA6700The Netherlands
3. Systematic Botany and Functional Biodiversity Institute of Biology Leipzig University Johannisallee 21‐23 Leipzig04103Germany
4. Department of Environmental Science and Ecological Engineering Korea University 145 Anamro Seongbukgu Seoul02841Korea