1. Department of Parks and Wildlife Science and Conservation Division 37 Wildlife Pl Woodvale Western Australia 6026 Australia
2. Institute for Applied Ecology University of Canberra Canberra Australian Capital Territory 2601 Australia
3. Australian Research Council Centre of Excellence for Coral Reef Studies James Cook University Townsville Queensland 4811 Australia
4. School of Life and Environmental Sciences University of Sydney Sydney New South Wales 2006 Australia