1. Geography Research Unit University of Oulu Oulu Finland
2. Ecology and Genetics Research Unit University of Oulu Oulu Finland
3. Department of Geosciences and Geography University of Helsinki Helsinki Finland
4. Department of Physiological Diversity Helmholtz Center for Environmental Research – UFZ Leipzig Germany
5. German Centre for Integrative Biodiversity Research (iDiv) Halle‐Jena‐Leipzig Leipzig Germany