1. Department of Biological Sciences Macquarie University North Ryde, Sydney New South Wales 2109 Australia
2. Sydney Institute of Marine Science Mosman New South Wales 2088 Australia
3. CSIRO Marine and Atmospheric Research Hobart Tasmania 7004 Australia
4. Department of Statistics and Institute for the Oceans & Fisheries University of British Columbia Vancouver British Columbia V1V 1V7 Canada
5. Institute for Marine and Antarctic Studies University of Tasmania Hobart Tasmania 7004 Australia