1. National Centre for Biological Sciences Tata Institute of Fundamental Research Bellary Rd Bangalore560065 India
2. School of Natural Resource Management Nelson Mandela University Madiba Rd George6529 South Africa
3. Rangeland Resources Research Unit U.S. Department of Agriculture, Agricultural Research Service Fort Collins Colorado USA
4. Scientific Services South African National Parks Skukuza South Africa
5. Faculty of Biological Sciences School of Biology University of Leeds Leeds UK