1. Department of Ecology & Evolutionary Biology University of Colorado Boulder Colorado 80309 USA
2. CU Museum of Natural History University of Colorado Boulder Colorado 80309 USA
3. Natural Resource Ecology Laboratory Colorado State University Fort Collins Colorado 80523 USA
4. Department of Natural Resources and the Environment University of New Hampshire Durham New Hampshire 03824 USA
5. Department of Computer Science University of New Hampshire Durham New Hampshire 03824 USA