1. State Key Laboratory of Biocontrol Guangdong Provincial Key Laboratory of Plant Resources, School of Life Sciences Sun Yat‐sen University Guangzhou 510275 China
2. State Key Laboratory of Palaeobiology and Stratigraphy Nanjing Institute of Geology and Palaeontology Chinese Academy of Sciences Nanjing 210008 China
3. School of Earth Science and Resources Chang’an University Xi’an 710054 China
4. Department of Botany and Plant Biotechnology University of Johannesburg Johannesburg 2006 South Africa
5. Komarov Botanical Institute of the Russian Academy of Sciences St Petersburg 197376 Russia