1. Montana Cooperative Fishery Research Unit, Department of Ecology Montana State University Bozeman Montana USA
2. U.S. Geological Survey, Montana Cooperative Fishery Research Unit Montana State University Bozeman Montana USA
3. Montana Fish, Wildlife, and Parks Great Falls Montana USA
4. Statistical Consulting and Research Services Montana State University Bozeman Montana USA
5. Department of Ecology Montana State University Bozeman Montana USA
6. U.S. Geological Survey Northern Rocky Mountain Science Center Bozeman Montana USA