1. National Oceanic and Atmospheric Administration, National Marine Fisheries Service, Northwest Fisheriest Science Center, Environmental Physiology Program Seattle Washington 98112 USA
2. Oregon Department of Fish and Wildlife Eastern Oregon University La Grande Oregon 97850 USA
3. Oregon Department of Fish and Wildlife Umatilla Hatchery Monitoring and Evaluation Umatilla Oregon 97882 USA
4. Oregon Department of Fish and Wildlife Bonneville Hatchery Cascade Locks Oregon 97014 USA