1. Department of Environment, Centre for Integrated Research, Conservation and Learning (CIRCLE); University of York; Wentworth Way, York United Kingdom
2. Flamingo Land Ltd., Kirby Misperton; North Yorkshire United Kingdom
3. Durrell Institute of Conservation and Ecology, School of Anthropology and Conservation; University of Kent; Canterbury, Kent United Kingdom
4. Department of Animal and Plant Sciences; University of Sheffield; Sheffield United Kingdom