1. Department of Genetics, Evolution and Environment University College London London UK
2. Centre for Biodiversity and Environment Research University College London London UK
3. Department of Animal and Plant Sciences NERC Biomolecular Analysis Facility University of Sheffield Sheffield UK
4. Department of Environmental Sciences Zoological Institute University of Basel Basel Switzerland
5. Department of Aquatic Sciences and Fisheries University of Dar es Salaam Dar es Salaam Tanzania