1. Biological and Environmental Sciences; University of Stirling; Stirling FK9 4LA United Kingdom
2. Department of Biological Sciences; Towson University; 8000 York Road Towson Maryland 21252 USA
3. Duke University Center for Tropical Conservation; Nicholas School of the Environment and Earth Sciences; PO Box 90318 Durham North Carolina 27708 USA
4. Florida Museum of Natural History; University of Florida; Gainesville Florida 32611 USA