1. School of Biological Sciences; University of Bristol; Woodland Road Clifton Bristol BS8 1UG UK
2. Botanisches Institut; Universität Bonn; Venusbergweg 23 Bonn D-53115 Germany
3. Department of Plant Cell Biology, Institute of Botany; Rheinische-Friedrich-Wilhelms-University of Bonn; Kirschallee 1 Bonn D-53115 Germany