1. Department of Microbial, Biochemical and Food Biotechnology University of the Free State Bloemfontein South Africa
2. Department of Animal Wildlife and Grassland Sciences University of the Free State Bloemfontein South Africa
3. Department of Zoology and Entomology University of the Free State Bloemfontein South Africa
4. Department of Genetics University of the Free State Bloemfontein South Africa