1. Graduate School of MedicineKyoto University 53 Shogoin-kawaharacho Sakyo-ku Kyoto 606-8507 Japan
2. Medical Sciences Innovation Hub Program, RIKEN Cluster for ScienceTechnology and Innovation Hub, Tsurumi-ku Kanagawa 230-0045 Japan
3. Foundation for Biomedical Research and Innovation at KobeCenter for Cluster Development and Coordination, Chuo-ku, Kobe Hyogo 650-0047 Japan