1. Key Laboratory of Alpine Ecology and BiodiversityInstitute of Tibetan Plateau Research, Chinese Academy of Sciences Beijing China
2. College of Resources and EnvironmentUniversity of Chinese Academy of Sciences Beijing China
3. State Key Laboratory of Systematic and Evolutionary BotanyInstitute of Botany, Chinese Academy of Sciences Beijing China
4. Estación Experimental de Zonas ÁridasConsejo Superior de Investigaciones Científicas Almería Spain