1. Department of Marine, Earth, and Atmospheric Sciences; North Carolina State University; Raleigh NC USA
2. Department of Geography; Loughborough University; Leicestershire UK
3. National Institute for Aquatic Resources; Danish Technical University; Lyngby Denmark
4. School of Biological Sciences; University of Essex; Essex UK
5. Now at The James Hutton Institute; Dundee UK
6. Now at School of Science and Technology; Nottingham Trent University; Nottingham UK