1. Hubrecht Institute-KNAW & University Medical Center; Utrecht The Netherlands
2. Department of Cell Biology; University Medical Center; Utrecht The Netherlands
3. Stem Cell Biology Group; Cancer Research UK Manchester Institute; The University of Manchester; UK
4. Laboratoire de Biologie du Développement; Sorbonne Universités; CNRS UMR7622; Inserm U 1156; IBPS; UPMC Univ Paris 06; France