1. Department of Forest Sciences; “Luiz de Queiroz” College of Agriculture; University of São Paulo; Pádua Dias Avenida 11 Piracicaba SP 13400-970 Brazil
2. Institute of Biology; University of Campinas - UNICAMP; Cidade Universitária Zeferino Vaz - Barão Geraldo; Campinas SP 13083-970 Brazil
3. Department of Ecology and Evolutionary Biology; University of Connecticut; Storrs Connecticut 06269-3043 USA