1. Division of Mucosal Immunology; Department of Microbiology and Immunology; The Institute of Medical Science; The University of Tokyo; Tokyo Japan
2. Core Research for Evolutional Science and Technology (CREST); Japan Science and Technology Agency (JST); Tokyo Japan
3. Division of Infectious Genetics; Department of Microbiology and Immunology; The Institute of Medical Science; The University of Tokyo; Tokyo Japan
4. Department of Microbiology and Immunology; Columbia University Medical Center; New York NY USA
5. International Research and Development Center for Mucosal Vaccine; The Institute of Medical Science; The University of Tokyo; Tokyo Japan