1. School of Environmental and Forest Services; University of Washington; 1325 Fourth Avenue, Suite 1515 Seattle WA 98101-2509 USA
2. School of Aquatic and Fishery Sciences; University of Washington; 1325 Fourth Avenue, Suite 1515 Seattle WA 98101-2509 USA
3. Missouri Department of Conservation; 3500 E Gans Road Columbia MO 65201 USA
4. Department of Fisheries and Wildlife Sciences; University of Missouri; 302 Natural Resources Building Columbia MO 65211 USA