1. College of Earth and Environmental Sciences, Lanzhou University Lanzhou PR China
2. Dunhuang Xihu National Nature Reserve Administration Jiuquan PR China
3. Centre for Invasion Biology, Department of Mathematical Sciences Stellenbosch University, African Institute for Mathematical Sciences Matieland South Africa
4. Key Laboratory of Desert and Desertification Northwest Institute Eco‐environment and Resource, Chinese Academy of Sciences Lanzhou PR China