1. US Geological Survey Columbia Environmental Research Center Columbia Missouri
2. Waterborne Environmental, Inc Leesburg Virginia USA
3. Present address: Department of Ecology, Evolution, and Organismal Biology Kennesaw State University Kennesaw Georgia USA
4. Present address: Department of Fisheries and Wildlife Michigan State University East Lansing Michigan USA
5. University of Maryland University College‐Asia Yokota Fussa‐shi Japan