1. Department of Natural Resource Ecology and Management; Oklahoma State University; Stillwater Oklahoma 74078 USA
2. U.S. Geological Survey; Oklahoma Cooperative Fish and Wildlife Research Unit; Department of Natural Resource Ecology and Management; Oklahoma State University; Stillwater Oklahoma 74078 USA
3. U.S. Fish and Wildlife Service; Southwestern Native Aquatic Resources and Recovery Center; Dexter New Mexico 88230 USA