1. Department of Animal and Plant Sciences; University of Sheffield; Sheffield S10 2TN UK
2. Evolution & Ecology Research Centre; School of Biological, Earth and Environmental Sciences; University of New South Wales; Sydney NSW 2052 Australia
3. Department of Zoology; University of Otago; PO Box 56 Dunedin 9054 New Zealand
4. Evolutionary Biology; Max Planck Institute for Ornithology; Seewiesen DE-82319 Germany