1. Institute of Life and Earth Sciences, School of Energy, Geoscience, Infrastructure and Society; Heriot-Watt University; Edinburgh United Kingdom
2. School of Marine Science and Engineering; Plymouth University; Plymouth United Kingdom
3. St. Abbs Marine Station; St. Abbs; Berwickshire United Kingdom
4. Department of Forestry, Wildlife and Fisheries and Center for Environmental Biotechnology; The University of Tennessee; Knoxville Tennessee USA