1. Institute of Transformative bio-Molecules, WPI-ITbM; Nagoya University; Furo-cho Chikusa-ku Nagoya 464-8602 Japan
2. Department of Radiation Genetics; Kyoto University Graduate School of Medicine; Sakyo, Yoshida-konoemachi Building D, 3F Kyoto 606-8501 Japan
3. Laboratory for Molecular Biosciences; Kyoto University Graduate School of Medicine; Yoshida-konoemachi Building E 606-8501 Kyoto Sakyo Japan