1. Department of Biological Sciences Royal Holloway University of London Egham TW20 0EX UK
2. Evolution and Ecology Research Centre School of Biological, Earth and Environmental Sciences University of New South Wales Sydney New South Wales Australia
3. The Carnivore Conservancy Ulverstone Tasmania Australia
4. Function, Evolution and Anatomy Research Laboratory School of Environmental and Rural Science University of New England Armidale New South Wales Australia
5. School of Life and Environmental Sciences The University of Sydney Sydney New South Wales Australia