1. Department of Physiology, Development and Neuroscience University of Cambridge Cambridge UK
2. MRC Weatherall Institute of Molecular Medicine, Radcliffe Department of Medicine University of Oxford Oxford UK
3. Department of Genetics University of Cambridge Cambridge UK
4. Division of Genetic Medicine, Department of Medicine Vanderbilt University School of Medicine Nashville Tennessee USA
5. Division of Molecular Neurobiology National Institute for Basic Biology Okazaki Japan