1. CAS Key Laboratory of Mountain Ecological Restoration and Bioresource Utilization, Chengdu Institute of Biology Chinese Academy of Sciences Chengdu China
2. University of Chinese Academy of Sciences Beijing 100049 China
3. State Key Laboratory of Biocatalysis and Enzyme Engineering, School of Life Sciences Hubei University Wuhan China
4. State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany Chinese Academy of Sciences Beijing China
5. College of Forestry Hainan University Haikou China