1. Molecular Invertebrate Systematics and Ecology Laboratory, Graduate School of Engineering and ScienceUniversity of the Ryukyus Nishihara Okinawa Japan
2. Dipartimento di Scienze della Vita e dell'AmbienteUniversità Politecnica delle Marche Ancona Italy
3. Tropical Biosphere Research CenterUniversity of the Ryukyus Nishihara Okinawa Japan