1. Department of Biological Sciences Macquarie University Sydney NSW 2109 Australia
2. Lancaster Environment Centre Lancaster University Lancaster LA1 4YQ U.K
3. Centre d'Ecologie Fonctionnelle et Evolutive CEFE UMR 5175 Campus CNRS 1919 Route de Mende F‐34293 Montpellier Cedex 5 France
4. School of Life Sciences University of Lincoln Riseholme Park Lincoln LN2 2LG U.K