1. Key Laboratory of Coastal Environmental Processes and Ecological Remediation Yantai Institute of Coastal Zone Research (YIC) Chinese Academy of Sciences (CAS) Shandong Provincial Key Laboratory of Coastal Environmental Processes, YICCAS Yantai Shandong 264003 P. R. China
2. University of Chinese Academy of Sciences Beijing 100049 P. R. China
3. Laboratory for Marine Biology and Biotechnology Pilot National Laboratory for Marine Science and Technology (Qingdao) Qingdao Shandong 266237 P. R. China
4. Center for Ocean Mega-Science Chinese Academy of Sciences Qingdao Shandong 266071 P. R. China