1. Chinese Academy of Geological Sciences Beijing China
2. State Key Laboratory of Palaeobiology and Stratigraphy Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences Nanjing China
3. College of Palaeontology Shenyang Normal University Shenyang China
4. Center for Excellence for Life Evolution and Environment Chinese Academy of Sciences Nanjing China
5. Shenyang Center of China Geological Survey Shenyang China
6. Institute of Geosciences, Division of Palaeontology University of Bonn Bonn Germany