1. State Key Joint Laboratory of Environment Simulation and Pollution Control School of Environment Tsinghua University Beijing 100084 China
2. Laboratory of Alpine Ecology and Biodiversity Institute of Tibetan Plateau Research Chinese Academy of Sciences Beijing 100085 China
3. Key Laboratory of Adaption and Evolution of Plateau Biota Northwest Institute of Plateau Biology Chinese Academy of Sciences Xining 810008 China
4. Shapotou Desert Experiment and Research Station Cold and Arid Regions and Environmental & Engineering Research Institute Chinese Academy of Sciences Lanzhou 730000 China
5. Chengdu Institute of Biology Chinese Academy of Sciences Chengdu 610041 China
6. Institute for Environmental Genomics and Department of Botany and Microbiology University of Oklahoma Norman Oklahoma 73019
7. Earth Sciences Division Lawrence Berkeley National Laboratory Berkeley California 94720