1. Laboratory of Ecology, Biology, and Physiology of Aquatic Organisms, Department of Biology, Tunis Faculty of Sciences University of Tunis El Manar Tunis Tunisia
2. Anatomy and Cytology Service CHU Mohamed Taher Maamouri Hospital Nabeul Tunisia
3. University of Carthage, Faculty of Sciences of Bizerte Laboratory of Environment Biomonitoring, Hydrobiology Unit Zarzouna Tunisia