1. State Key Laboratory of Vegetation and Environmental Change, Institute of BotanyChinese Academy of Sciences Beijing China
2. College of Resources and EnvironmentUniversity of Chinese Academy of Sciences Beijing China
3. Key Laboratory of Adaptation and Evolution of Plateau Biota, Northwest Institute of Plateau BiologyChinese Academy of Sciences Xining China
4. Department of Ecology, College of Urban and Environmental SciencesPeking University Beijing China
5. Key Laboratory of Ecosystem Network Observation and Modeling, Institute of Geographic Sciences and Natural Resources ResearchChinese Academy of Sciences Beijing China
6. Key Laboratory of Forest Ecology and Management, Institute of Applied EcologyChinese Academy of Sciences Shenyang China