1. Global Change and Sustainability Center; University of Utah; Salt Lake City Utah USA
2. Department of Ecology, Evolution, and Organismal Biology; Iowa State University; Ames Iowa USA
3. Department of Biology and the Ecology Center; Utah State University; Logan Utah USA
4. Department of Geology and Geophysics; University of Utah; Salt Lake City Utah USA
5. Department of Biology; University of Utah; Salt Lake City Utah USA
6. Department of Earth System Science; University of California; Irvine California USA