1. Department of Botany and Plant Biotechnology; African Centre for DNA Barcoding; University of Johannesburg; APK Campus, PO Box 524 2006 Johannesburg South Africa
2. Department of Organismic and Evolutionary Biology; Harvard University; Cambridge Massachusetts 02138 USA
3. McGill University; 1205 Avenue du Docteur-Penfield Montreal Quebec H3A 1B1 Canada
4. Quebec Center for Biodiversity Science; McGill University; 1205 Avenue du Docteur-Penfield Montreal Quebec H3A 1B1 Canada
5. Department of Geography, Environmental Management and Energy Studies; University of Johannesburg; APK Campus 2006 Johannesburg South Africa