1. Wildlife Conservation Research Unit, Department of Zoology, University of Oxford, Recanati‐Kaplan Centre, Tubney House, Abingdon Road, Tubney, Abingdon, Oxfordshire OX13 5QL, United Kingdom
2. NERC Biomolecular Analysis Facility, Department of Animal and Plant Sciences, University of Sheffield, Western Bank, Sheffield, South Yorkshire, S10 2TN, United Kingdom
3. Behavioural Ecology and Self‐Organization, University of Groningen, P.O. Box 11103, 9700 CC Groningen, The Netherlands
4. Theoretical Biology, University of Groningen, P.O. Box 11103, 9700 CC Groningen, The Netherlands