1. Faculty of Life and Medical Sciences; Doshisha University; Tatara, Miyakotani, Kyotanabe Kyoto 610-0321 Japan
2. Research Organization of Science and Technology; Ritsumeikan University; Nogihigashi, Kusatsu Shiga 525-8577 Japan
3. Division of Biological Science; Graduate School of Science; Nagoya University; Furo-cyo, Chikusa-ku, Nagoya Aichi 464-8602 Japan