1. Department of Ecology and Evolutionary Biology; University of Tennessee; 569 Dabney Hall 1416 Circle Drive Knoxville Tennessee 37996 USA
2. Center for Macroecology, Evolution and Climate; Natural History Museum of Denmark; University of Copenhagen; Sølvgade 83S DK-1307 Copenhagen K Denmark
3. Department of Zoology; University of Wisconsin-Madison; Madison Wisconsin 53706 USA
4. Department of Entomology; University of Wisconsin-Madison; Madison Wisconsin 53706 USA