1. Animal Evolution and Systematics Group (AES); Department of Biological Sciences; University of Cape Town; Cape Town South Africa
2. Department of Biology; University of Massachusetts Amherst; 221 Morrill Science Center Amherst MA 01003 USA
3. Centre for Statistics in Ecology, Environment and Conservation (SEEC); Department of Statistical Sciences; University of Cape Town; Cape Town South Africa
4. South African National Biodiversity Institute (SANBI); Cape Town South Africa