1. Department of Wildlife, Fisheries and Conservation Biology; University of Maine; 5755 Nutting Hall Orono ME 04469 USA
2. Department of Environmental Conservation; University of Massachusetts, Amherst; 160 Holdsworth Way Amherst MA 01003 USA
3. U.S. Geological Survey; Maine Cooperative Fish and Wildlife Research Unit; Orono ME USA
4. U.S. Geological Survey; West Virginia Cooperative Fish and Wildlife Research Unit; West Virginia University; Morgantown WV USA