1. Wildlife Biology Program, Department of Ecosystem and Conservation Sciences; W.A. Franke College of Forestry and Conservation, University of Montana; 32 Campus Drive Missoula MT 59812 USA
2. Alberta Conservation Association; 817 4th Avenue South #400 Lethbridge AB T1J 0P3 Canada
3. Wildlife Biology Program, Department of Ecosystem and Conservation Sciences; W.A. Franke College of Forestry and Conservation, University of Montana; Missoula, 32 Campus Drive Missoula MT 59812 USA