1. Department of Organismic and Evolutionary Biology and Harvard University Herbaria Harvard University Cambridge Massachusetts 02138 USA
2. Délégation à la Recherche Government of French Polynesia B.P. 20981 Papeete, Tahiti French Polynesia
3. Association Te Rau Ati Ati a Tau a Hiti Noa Tu B.P. 11553 Mahina, Tahiti French Polynesia