1. School of Earth Sciences University of Bristol Life Sciences Building, Tyndall Avenue Bristol BS8 UK
2. State Key Laboratory of Palaeobiology and Stratigraphy Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment Chinese Academy of Sciences Nanjing 210008 China
3. CNRS, Géosciences Rennes, UMR 6118 Université de Rennes Rennes 35000 France